General 
					 
					There are about 
					70 species of butterworts worldwide. Many 
					are found in 
					Mexico and in Europe.
					
					 They grow on wet, dripping, 
					moss-covered rocks in the mountains, on moist, grass-covered 
					sandy ditches in savannas, and in other similar conditions, 
					sometimes in company of other carnivorous plants. Of six 
					species found in the southeastern United States, five are 
					indigenous to the region. Three other species grow in 
					northern U.S. and Alaska. 
					
					Butterworts are a rosette of thin leaves, 
					typically lying prostrate, with white, fibrous roots. The 
					upper surface of the leaf is covered with short stalked 
					glands that secrete clear, sticky mucilage to trap small  
					animal prey. The leaf feels greasy to the touch because of 
					this secretions. This gave the plant its common name. The 
					genus name Pinguicula is derived from the Latin 
					word pinguis, meaning fat. 
					
					
					 Many butterwort rosettes have close 
					resemblance to each other, and it is often difficult to 
					distinguish one species from another when the plants are out 
					of flower. The leaf is yellowish-green in most species but 
					has a distinctive reddish color in some species (including 
					one U.S. species). The diameter of the rosette varies 
					anywhere from 2  to 15 cm depending on the species. 
					  
					
					
					  
					
					Prey Capture and Digestion 
					
					Most of the prey 
					the butterworts are capable of trapping is small  insects such 
					as gnats, mosquitoes 
					and small  ants. When an insect lands or crawls on 
					the upper surface of the butterwort's leaf, the insect is mired down to the 
					surface by the sticky substance secreted by the stalked 
					glands, as in sundews. Unlike sundew tentacles, however, 
					these short stalked glands offer no movement. The stalked 
					gland consists of 16 radiating glandular cells supported on 
					a single-celled stalk. 
					
					
					 Scattered over 
					much of the leaf surface 
					are another kind of glands with a similar structure. These 
					are sessile (stalkless) glands consisting of 8 radiating 
					glandular cells. These sessile glands are responsible for the secretions  of 
					digestive fluids and the subsequent absorption of the 
					products of digestion. Studies have shown that nitrogenous 
					compounds placed on the leaf surface precipitate the enzyme
					secretions  from sessile 
					glands in a matter of minutes.  
					When a small  insect -- or any 
					nitrogenous substance such as a piece of meat -- is placed 
					on the leaf surface, there is copious secretions  of digestive 
					fluid from the sessile glands. Often a trapped insect 
					sinks down to the leaf surface -- completely submerged in 
					the digestive fluids -- and comes 
					into direct and tight contact with sessile glands on the 
					leaf surface. The secretions are also shown to contain an 
					antiseptic substance which effectively prevents bacterial 
					activities during the course of the digestive process if the 
					prey is small  enough which, in nature, is usually the case. 
					
					The resultant digested material is 
					absorbed through the sessile glands. Studies using 
					radioactive isotope (carbon 14) show that products of 
					digestion are rapidly taken into the leaf in a matter of a 
					few hours and are carried to the other parts of the 
					plants. 
					
					Leaf Movement 
					
					Often the leaf movement is observed in 
					association with the capture of prey. When a small  prey is 
					placed on the leaf surface near the leaf margin, the leaf 
					begins to curl up. 
					
					 It is a slow motion taking several hours 
					and possibly lasting several days. Also, the amount of 
					curling is rather limited. It is unlikely, therefore, that 
					the leaf movement contributes to the entrapment of the prey 
					in the same degree as observed in the leaf-folding in some 
					sundew species. 
					It is also noted that the contact of an 
					insect with the leaf (away from the margin) results in the 
					dishing of the leaf area below the prey.  
					
					All leaf movements are known to be 
					caused by the stimulation to the leaf surface and are 
					attributed to a growth phenomenon. Although not as active as 
					tentacle bending or leaf folding in sundews, the leaf 
					movement in butterworts is a matter of common observation. 
					How much significance should be given 
					to it is debatable. A general 
					interpretation is that the upwardly curled leaf 
					margin -- and localized dishing effect
					of the leaf by stimulation 
					-- help to hold the secretions  in place during the digestion 
					process.  
					
					Inflorescence 
					
					Butterworts produce a showy 
					display of colorful flowers which are borne 
					singly (for most species) at the tip of a 
					slender, often glandular, pubescent scape. Early spring is 
					the flowering season for most U.S. species. 
					
					 The zygomorphic 
					(laterally symmetric) flower has a sympetalous (united 
					petals) corolla forming a cylindrical tube which divides 
					into five lobes (2 lobes on the upper lip 
					& 3 on the lower lip) on the front and terminates in a slender 
					spur on the back.  The spur is a nectar 
					container, a structure commonly seen in insect-pollinating 
					flowers. Near the lower entrance of the corolla tube 
					projects a hairy structure known as a palate. The 
					microscopic structure of the palate hairs, along with the 
					color and shape of the entire corolla, is characteristic to 
					each species and can be used for the identification. On the upper side of the inner corolla tube are 
					located two stamens and a stigma lobe. A wide stigma lobe 
					covers anthers for incoming pollinators, thus providing a 
					mechanism to discourage self-pollination. 
					After the flower, the scape grows 
					further as it straightens itself. The seeds mature in a 
					month or so. The seed capsule contains hundreds of tiny, 
					powder-like seeds. Seeds have a distinctive surface pattern 
					for each species. 
					
					
					  
					
					Uses 
					
					Apart from the carnivorous nature of 
					the butterwort plants, people in Scandinavian countries have 
					mixed the leaf extracts with fresh milk to make it curdle. 
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